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Submission Preparation Checklist

As part of the submission process, authors are required to check off their submission's compliance with all of the following items, and submissions may be returned to authors that do not adhere to these guidelines.
  • The submission has not been previously published, nor is it before another journal for consideration (or an explanation has been provided in Comments to the Editor).
  • The manuscript is in ENGLISH.
  • The submission file is in OpenOffice, Microsoft Word, or RTF document file format.
  • All authors’ ORCID ID were included.
  • Where available, URLs for the references have been provided.
  • The text is double-spaced between lines, justified text, and Calibri font, body 11 (eleven) (except for special symbols not included in this font).
  • Pictures, graphics and illustrations even inserted in the text must necessarily be sent in separate files, in the original format in which they were created.
  • The text adheres to the stylistic and bibliographic requirements outlined in the Author Guidelines.
  • Authors declare that there is no conflict of interest; otherwise, it must be specified in “Comments to the Editor”.
  • An authors’ contributions statement must be included in a section at the end of the article for works written by more than 4 (four) authors. The statement has to include the individual contribution of each author.

Author Guidelines


IMPORTANT: You must correctly complete all fields for each AUTHOR: Name, Middle Name, Last Name, Email, ORCID, URL (if there is), Institution/Affiliation, Country, and Biography Summary.

From 2019 on, the inclusion of the ORCID ID for each author is mandatory.

The fields for the manuscript should also be filled with the information: Title, Abstract, Area and Sub-Area of Knowledge, Keywords, Language and Funding/Support Agencies.

  • The manuscript must be sent only in ENGLISH.
  • Manuscripts should be sent in the format “.DOCX” or “.RTF” to the journal submission system, along with a submission letter explaining the importance and originality of the study.
  • All submissions are performed through the “Portal de Revistas da USP”.
  • Pictures, graphics, and illustrations even inserted in the text must necessarily be sent in separate files, in the original format in which they were created.
  • The files must be sent in separate files formats: “.PSD”, “.TIF”, “.JPG” “.PCX”, “.GIF” or “.BMP” for figures; “.EPS”, “.CDR”, “.WMF”, “.AI”, “.PPTX” or “.XLSX” for graphics; and “.MOV”, “.AVI”, “.MPG”, “.MP4” or “.M4V” for videos.
  • Tables should be placed at the end of the manuscript.
  • Always keep updated the email and the address of the author(s) or corresponding.
  • The authors may suggest potential reviewers.



  • The manuscripts must be double-spaced between lines, justified text, and Calibri font, body 11 (eleven) (except for special symbols not included in this font).
  • The text should be organized in the following sections, each started on a new page, in the order, and numbered consecutively: Title Page, Abstract with Keywords, Body Text, Acknowledgments, References, Appendices, Tables, and Figure Legends.
  • Scientific names of species and genera, and other Latin terms, must be italicized in all sections of the manuscript.

(1) TITLE PAGE: Should include the Title, Running Title, Author(s) Name(s), ORCID numbers(s), Institution(s), and Address(es). The title should be concise and, where appropriate, should include particulars about families and/or taxa of higher categories. New taxa names should not be included in the titles.

(2) ABSTRACT WITH KEYWORDS: All papers should have an abstract and keywords in English. The quality of the abstract is of great importance since it can be reproduced in other vehicles. Therefore, must be written in an intelligible form as it may be published separately, and should summarize the main facts, ideas, and conclusions of the article. Telegraphic abstracts are unacceptable. Finally, the abstract should include all new taxonomic names for reference purposes. Abbreviations should be avoided. It should not include bibliographical references. Abstracts and keywords must not exceed 350 (three hundred and fifty) and 5 (five) words, respectively.

(3) BODY TEXT: The main body text varies with different types of papers, but should usually include the following sections: Introduction, Material and Methods, Results, Discussion, Conclusion, Acknowledgments, and References. The main headings of each section of the body of the text should be written in capital letters, bold and centered. Secondary headings should have only the first letter capitalized, bolded, and centered. Tertiary headings should be the first letter capitalized, in bold and indented left. In all cases, the text should start on the next line.

(4) REFERENCES: Citations in the text should be given as follows: Martins (1959) or (Martins, 1959) or (Martins, 1959, 1968, 2015) or Martins (1959a, b) or Martins (1959: 14-20) or Martins (1959: figs. 1, 2) or Martins & Reichardt (1964) or (Lane, 1940; Martins & Chemsak, 1966a, b) or Martins et al. (1966) or (Martins et al., 1966), the latter when the article contains 3 (three) or more authors. There is no need to provide a full reference when authors and date are presented only as an authority of taxa.

(5) REFERENCES MODELS: References should be arranged alphabetically and according to the following format, respecting the titles in italics. Journal titles must be written in full (not abbreviated):

  • Journal Article - Author(s). Year. Article title. Journal name, volume: initial page-final page.
  • Journal Article (print and online version with differents dates, for instance: ahead-of-print, online first, etc.) - Author(s). Year online. Article title. Journal name, volume: initial page-final page, Year on print.
  • Books - Author(s). Year. Book title. Publisher, Place of publication.
  • Books Chapters - Author(s). Year. Chapter title or part. In: Author(s) or Editor(s), Book title. Publisher, Place of publication, volume or part, initial page-final page.
  • Dissertations and Theses - Author(s). Year. Title of dissertation or thesis. (Masters Dissertation or Doctoral Thesis). University, Place of publication.
  • Electronic Publications - Author(s). Year. Document title. Available at: electronic address. Accessed: DD/MM/AAAA.
  • Datasets - Author(s). Year. Title. Version. Publisher. [dataset]. Available at: (preferably the DOI number as url). Accessed: DD/MM/AAAA. - See more in the Research Data section.


  • All tables should be numbered in the same sequence in which they are cited in the text.
  • Legends should be self-explanatory, without the need to resort to text.
  • Tables should be formatted preferentially towards portrait, remaining the sense landscape for exceptional cases.
  • In the text, tables should be referred as Table 1, Tables 2 and 4, Tables 2-6.
  • Use “TABLE” in the table(s) heading(s).


  • All figures should be numbered in the same sequence in which they are cited in the text.
  • Each illustration of a composite figure should be identified by a capital letter and referred to in the text as, for example: Fig. 1A, Fig. 1B.
  • When possible, the letters must be positioned in the lower-left corner of each illustration of a composite figure.
  • Photographs in black and white or color must be sending at high resolution (300 DPI minimum).
  • Use “Fig(s).” for referring to figures in the text, and “fig(s).” when referring to figures in another paper.
  • Use “FIGURE” in the figure(s) heading(s).
  • The figures will be printed in black and white but maintained in color in the final PDF file.


  • Page-proofs with the revised version will be sent by email to the corresponding author.
  • Page-proofs must be returned to the Editor, preferentially within 48 (forty-eight) hours.
  • Only necessary corrections in proof will be permitted.
  • The omission of return the proof will mean automatic approval of version with no changes and may result in a delay in publication.


  • There is no publication fee to submit or publish in Papéis Avulsos de Zoologia.


  • Previously published content as figures, tables, illustrations, etc. must be accompanied by written permission from the copyright holder to reproduce.


  • The scientific content and opinions expressed in this publication are sole responsibility of the respective authors.


All Authors must meet the two minimum criteria:

  • Actively participate of the results discussion.
  • Review and approve the final version of the paper.

An authors' contributions statement must be included in a section at the end of the article for works written by more than 4 (four) authors. 

The statement has to include the individual contribution of each author.

Acknowledgments section may contain a declaration of other kinds of contributions.



  • Author must declare if there is any conflict of interest at the submission of the manuscript.
  • If there is no conflict of interest, the authors should select in the submission form the option: “Authors declare that there is no conflict of interest”; otherwise, it must be specified in “Comments to the Editor”.
  • Coeditors, Associated Editors, and Reviewers must declare to the Editor any conflict of interest before starting the review process.
  • For further information on Conflict of Interest refer to Council of Science Editors White Paper on Publication Ethics (English or Portuguese versions).


  • Research grants from funding agencies (please inform the research funder and the grant number) or any financial support must be declared.


  • Papéis Avulsos de Zoologia recommends the deposit of scientific data in appropriate repositories and its citation in accordance with FAIR principles (Findable, Accessible, Interoperable and Re-usable).
  • As described in the “General Information” section of this Instructions to Authors, all nucleotide sequence data (aligned as well as unaligned) should be submitted to GenBank or European Molecular Biology Laboratory (EMBL). All articles that contain nomenclatural acts are registered in Zoobank by the journal staff and the Life Science Identifier (LSID) of the article is included in the published version.
  • Authors are encouraged to deposit their data in a repository that is best suited to their dataset. Some examples of repositories are the Global Biodiversity Information Facility (GBIF), Environmental Data Initiative (EDI), Dryad, Figshare, Zenodo, etc.
  • The data repository used must guarantee the preservation of the data and provide a persistent identifier such as a DOI so it can be accessible and citable.
  • Journal will make exceptions for sensitive information as the location of endangered species.
  • Datasets used in the research, deposited in a scientific research data repository, should be cited in the “Materials and method section” and its reference (with the DOI number) should be included in the reference list.

Datasets References Examples:

  • Dataset deposited in a scientific research data repository:

    Botham, M.; Roy, D.; Brereton, T.; Middlebrook, I.; Randle, Z. 2013. United Kingdom Butterfly Monitoring Scheme: species trends 2012. NERC Environmental Information Data Centre. [dataset]. Available at: https://doi.org/10.5285/5afbbd36-2c63-4aa1-8177-695bed98d7a9 . Accessed: 13/04/2019.

    United States Department of Health and Human Services. Substance Abuse and Mental Health Services Administration. Office of Applied Studies. 2015. Treatment Episode Data Set -- Discharges (TEDS-D) -- Concatenated, 2006 to 2011. Version V5. Ann Arbor, MI: Inter-university Consortium for Political and Social Research [distributor], 23 Nov. 2015. Available at: https://doi.org/10.3886/ICPSR30122.v5 . Accessed: 30/09/2019. 
  • Article supplementary information deposited in a scientific research data repository:

    Yoon, J; Sofaer, H.R, Sillet, T. S, Morrison, S.A., Ghalambor, C.K. 2017. Data from: The relationship between female brooding and male nestling provisioning: does climate underlie geographic variation in sex roles?. Journal of Avian Biology, 48(2):220-228. Available at: https://doi.org/10.5285/5afbbd36-2c63-4aa1-8177-695bed98d7a9 . Accessed: 07/07/2019.
  • Dataset published as data paper:
    Bovendorp, R. S., Villar, N., de Abreu-Junior, E. F., Bello, C., Regolin, A. L., Percequillo, A. R., & Galetti, M. 2017. Atlantic small-mammal: a dataset of communities of rodents and marsupials of the Atlantic forests of South America. Ecology, 98(8):2226–2226. [data paper]. Available at: http://dx.doi.org/10.1002/ecy.1893. Accessed: 07/10/2019.


  • Authors can share the published version¹ of the manuscript with their colleagues and post them on personal websites or institutional repositories for academic purposes while providing bibliographic details that credit, if applicable, its publication in this journal.
    ¹ Published version - is the article published in PDF format available at the Papéis Avulsos de Zoologia website.


  • Research or publication misconduct (plagiarism, self-plagiarism, fabrication or falsification of data and results, etc.) will be treated in accordance with the Commission on Publication Ethics (COPE) guidelines.
  • From May 2020 on, the journal has adopted the tool Originality Check (Turnitin) to prevent duplication and plagiarism occurrences.
  • Allegations, corrections, and doubts must be sent to the Editor (contact information). All allegations/cases will be evaluated and, when necessary, retractions, corrections or expressions of concern will be published.


For more details of the manuscript preparation format, see CBE Style Manual, available at the electronic address of the Council of Science Editors.

Papéis Avulsos de Zoologia is a publication of the Museu de Zoologia da Universidade de São Paulo.

Always consult the instructions to authors updated on the electronic pages: www.scielo.br/paz or www.revistas.usp.br/paz.

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