Catalogue of Bibionidae ( Diptera : Bibionomorpha ) types housed in the collection of the Museu de Zoologia da Universidade de São Paulo , Brazil

Following a recommendation of the International Code of Zoological Nomenclature, this paper provides a catalogue of the type specimens of Bibionidae (Diptera: Bibionomorpha) held in the collection of the Museu de Zoologia da Universidade de São Paulo, Brazil (MZUSP). Label data and the condition of 21 type specimens (two holotypes and 19 paratypes) of two Neotropical species is provided. Photographs of the male terminalia of the holotypes are also presented. Key-Words. Nomenclature; Scientific collections; Taxonomy; Type material.


INTRODUCTION
Brazil is one of the most diverse countries in the world and its most important scientific collection is the Museu de Zoologia da Universidade de São Paulo (MZUSP), São Paulo, Brazil.This collection includes more than 8,000 species and six hundred thousand mounted specimens, being the largest and most important depository of Neotropical Diptera in Brazil and in South America.Additional information on the collection is available at www. mz.usp.br.
Following a recommendation of the International Code of Zoological Nomenclature (ICZN, 1999: 72F.4), the present paper provides a catalogue with information on 21 type specimens (two holotypes and 19 paratypes) of two Neotropical species of the family Bibionidae, genus Bibionellus Edwards, held in the collection of the MZUSP (Table 1).

MATERIALS AND METHODS
The list given below is in alphabetical order of subfamily, genus, and epithet.Names, combinations, and spellings follow the catalogue of Bibionidae of the Americas south of the United States (Hardy, 1966).The standard for reproducing the information on the labels follows previous catalogues of general Diptera and Bibionomorpha housed in other major museums collections of Neotropical fauna (e.g., for the Museu de Zoologia da Universidade de São Paulo (MZUSP), Brazil (Fachin & Lamas, 2015;Falaschi et al., 2016b;Mello & Lamas, 2011)).All types listed here are drypinned specimens.High-resolution photographs will be subsequently provided for most primary types at MZUSP's homepage (www.mz.usp.br).(male genitalia).Ref. Hardy, 1953: 344 (identification key).
Type locality: Brazil, Goiás, Corumbá de Goiás.We analyzed the specimen and, in spite of some intraspecific color variation, it is a male specimen belonging to the species B. paulistensis.Hence, even in the absence of the identification label, we listed it together with the type-series of B. paulistensis and added a new identification label (label 4 previously listed).Our decision was based on the analysis of the shape of the parameres and the mesosome (including its lateral lobes and median and lateral projections), the presence of setae at the base of the gonocoxite, and the absence of long setae at the apex of the gonocoxite (Figs. 4 and 6).It is worth of note that a revision and phylogeny of the genus Bibionellus, including an identification key and illustration of all Bibionellus species known was done by Pinto & Amorim (1997).

REMARKS
According to Lane & Forattini (1948: 571), the holotype and nine paratypes of B. barrettoi, including the allotype, were deposited at the "Faculdade de Saúde Pública" (FSP-USP) and the remainder of the type-series were housed at the Museu de Zoologia (MZUSP).Lane & Forattini did the same with the type-series of B. paulistensis, establishing that the holotype, allotype, and two paratypes would be deposited at the FSP-USP and the remainder of the type-series would be housed at the MZUSP (Lane & Forattini, 1948: 573).However, since the beginning of the 1970s, the complete type-series is deposited at the MZUSP.
John Lane was a professor at the Department of Parasitology of the "Faculdade de Saúde Pública" (School of Public Health) of Universidade de São Paulo where he started the Laboratory of Entomology and an insect collection focused on Culicidae and other nematocerous flies.In 1970, the Department of Parasitology was transferred to the Institute of Biomedical Sciences and the entomological collection of John Lane was moved to the Museu de Zoologia (MZUSP), except for the Culicidae and other families of nematocerous flies of medical importance, which were kept in the Faculdade de Saúde Pública.Hence, the MZUSP is the depository of the types previously located at the FSP-USP.

Table 1 .
Type specimens of Bibionidae held in the MZUSP, in alphabetical order of species epithet, with reference to the number of specimens, males, and females, for each type category.
This paratype did not originally have an identification label.According to Lane & Forattini 1948: 571 there is a paratype from Porto Cabral, state of São Paulo, collected by Travassos Filho, Carrera, and Dente, in x.1941.