New species of Cerambycinae (Coleoptera, Cerambycidae) from Guatemala

. Three new species from Guatemala are described and illustrated: Ectenessa canoi sp. nov. (Ectenessini), from Petén; Oxycoleus schusteri sp. nov. (Oxycoleini), from Izabal; and Odontocera nigroaurantia sp. nov. (Rhinotragini), from Petén. Distinguishing characters from closely related species are provided.


INTRODUCTION
Guatemala has a complex topography and vegetation, which is reflected in its great biodiversity (Cano, 2006).In this country, the diversity of Cerambycidae was first summarized by Hovore (2006), who reported 769 species.Currently, Tavakilian & Chevillotte (2021), listed six subfamilies, 72 tribes, 350 genera, and 894 species and subspecies of cerambycids from Guatemala.
During the research stay of the first author in the University of the Valley of Guatemala (UVG) several unidentified coleopteran specimens were studied including those that represent undescribed species, which are described in the present work.

MATERIAL AND METHODS
Morphological structures were studied using a Zeiss Stemi 305 stereomicroscope (Carl Zeiss, Jena, Germany).Measurements and pictures were taken with a Zeiss® AxioZoom V16 stereomicroscope, with a Zeiss® AxioCam MRc5, 5 megapixels camera, controlled by ZEN (Zeiss Efficient Navigation) application.
References and classification of known species are restricted to their original description, information in catalog by Monné (2022) and checklist by Bezark (2022a).The acronyms used in the text is as follows: UVGC University of the Valley of Guatemala Collection, Guatemala, Guatemala.
Etymology: This species is dedicated to the recently deceased Enio Cano (Universidad del Valle de Guatemala), for his friendship, enthusiasm, and contribution to the knowledge of the diversity and conservation of Passalidae and Scarabaeidae in Guatemala.
Head: Frons narrow, transverse, densely microscopically punctate, with fine, sparse punctures interspersed; with minute orangish-brown pubescence not obscuring integument, and orangish-brown long, erect setae laterally.Median groove deep, somewhat wide from clypeus to area between antennal tubercles.Vertex and area between antennal tubercles and upper eye lobes with coarse, shallow, sparse punctures, punctures sparser close to prothorax; with minute orangish-brown pubescence not obscuring integument, light orange pubescence on orange area; with long, erect, scattered orangish-brown setae; with longitudinal gibbosity on median groove, from frons to orange area.Area behind lower eye lobes transversely striate, except smooth inferior area; striate area with short, orangish-brown setae; smooth area almost glabrous.Genae with transverse row of minute punctures near eye, smooth on remaining surface; with minute, decumbent bright orange setae on punctate area, glabrous on remaining surface.Central area of postclypeus microscopically punctate, except coarsely, sparsely punctures close to anteclypeus; with minute orangish-brown pubescence.Sides of postclypeus smooth and glabrous.Labrum with ternally and posteriorly with narrow semicircular sulcus; with coarse, shallow sparse punctures, denser and finer on gibbosities; with minute, dense bright orange pubescence not obscuring integument, somewhat darker on lateral gibbosities; apex of antero-and posterocentral gibbosities glabrous; with long, erect, scattered orangish setae.Sides of prothorax with sculpturing and pubescence as on pronotum.Prosternum finely, transversely striate; with short, erect orangish-yellow setae centrally close to coxal cavities and laterally, absent on remaining surface.Prosternal process very narrow on anterior half, strongly widened posteriorly, with apex truncate; with short, erect bright orange setae.Mesoventrite with short, erect orangish-yellow pubescence, not obscuring integument; mesanepisternum and mesepimeron with dense, minute bright orange pubescence not obscuring integument.Metanepisternum and sides of metaventrite with sparse orangish-brown pubescence not obscuring integument, and long, erect orangish-yellow setae interspersed, almost glabrous on central area of metaventrite.Scutellum glabrous, except narrow fringe of short orangish-yellow setae along margins.Elytra: Reaching about apex of abdominal segment II; slightly narrowed toward apex, distinctly dehiscent internally from anterior third; lateral carina well-marked, especially from anterior third; coarsely and abundantly punctate; with orangish-brown pubescence not obscuring integument; with a few short, erect orangish-yellow setae; apex rounded.Legs: Coxae and trochanters with short, sparse orangish-yellow setae.Profemora fusiform, meso-and metafemora pedunculate; with short, erect, somewhat orangish-brown setae; with long, erect, sparse brown setae.Tibiae with short, erect orangish-brown setae, and long, erect brown setae interspersed, erect setae distinctly denser dorsally and ventrally.Metatarsomere I longer than II-III together.
Etymology: This species is named after Jack Schuster (Universidad del Valle de Guatemala), a prominent entomologist interested in the study of the diversity and conservation of Guatemala, and author of several publications on the family Passalidae.
Head: Frons moderately coarsely, abundantly punctate, with yellowish-white pubescence.Median groove distinct from postclypeus to vertex, especially between eyes.Vertex and area behind upper eye lobes coarsely, abundantly punctate; both short and long yellowish setae, sparser behind upper eye lobes.Antennal tubercles elevated, somewhat coarsely, shallowly, sparsely punctate; with short, erect, scattered yellowish setae.Area behind lower eye lobes shallowly rugose-punctate; with short, sparse yellowish setae, interspersed with long, erect yellowish setae close to eye.Genae about half length of lower eye lobe; moderately finely, sparsely punctate except smooth apex; with short, sparse yellowish setae except glabrous smooth area, with long, erect setae of same color interspersed toward ventral surface.Wide central area of postclypeus with sculpturing as on frons, except smooth sides; with yellowish-white pubescence partially obscuring integument centrally, glabrous laterally; with one long, erect, yellowish seta on each side.Gulamentum smooth, glabrous on posterocentral anterior third, followed by narrow, smooth band; with abundant, long, erect yellowish setae.Narrowest area of prosternal process 0.25 times width of procoxal cavity; posterior area triangular.Mesoventrite finely, densely punctate, somewhat striate; with abundant yellowish pubescence, slightly denser laterally.Mesanepisternum and mesepimeron with dense, thick yellowish pubescence.Metaventrite and metanepisternum with dense yellowish-white pubescence, denser and thicker on anterior half of metaventrite and laterally on metanepisternum; with long, erect, sparse yellowish setae, denser laterally on metaventrite.Scutellum with short, scattered yellowish setae, apex emarginate.Elytra: Coarsely, abundantly punctate, punctures slightly finer toward apex, sparser on vitreous area; sides longitudinally concave from near base to near apex; sutural margins slightly divergent from posterior half; with very short, erect yellowish setae, gradually sparser toward apex; with long, erect, sparse yellowish setae on anterocentral area, close to scutellum; vitreous area with short, sparse yellowish setae; apex obliquely truncate, with short, erect yellowish setae.Legs: Coxae and trochanters with long, erect yellowish-orange setae.Femora with long, erect, sparse yellowish setae.Protibiae with short, erect yellowish-brown setae, distinctly denser ventrally, and long, erect setae on outer margin; meso-and metatibiae with long, erect, sparse yellowish setae, slightly denser laterally and ventrally.Tarsi with long, erect brownish setae.Metatarsomere I as long as II-III together.

Etymology:
The specific name nigroaurantia is composed from the Latin "niger, nigro" (black) and "aurantia" (orange), referring to the combination of black and orange integument.
: OP-F: Conceptualization, Methodology, Writing -original draft; AS-S: Writingreview & editing; OP-F, AS-S: Investigation.All authors actively participated in the discussion of the results; they reviewed and approved the final version of the paper.CONFLICTS OF INTEREST: Authors declare there are no conflicts of interest.FUNDING INFORMATION: Programa de Apoyo para Estudios de Posgrado (PAEP), Program in Biological Sciences of the UNAM for funding travel to the University of the Valley of Guatemala Collection.