Barrio-Amorós, C.L. et al. 2010. An addition to the diversity of dendrobatid frogs in Venezuela: description of three new collared frogs (Anura: Dendrobatidae: Mannophryne). Phyllomedusa: Journal of Herpetology. 9, 1 (Jul. 2010), 03–35. DOI:https://doi.org/10.11606/issn.2316-9079.v9i1p03-35.