Lindquist, E. et al. 2012. Annular bone growth in phalanges of five Neotropical Harlequin Frogs (Anura: Bufonidae: Atelopus). Phyllomedusa: Journal of Herpetology. 11, 2 (Dec. 2012), 117–124. DOI:https://doi.org/10.11606/issn.2316-9079.v11i2p117-124.