Duellman, W. E., Jungfer, K.-H., & Blackburn, D. C. (2011). The phylogenetic relationship of geographically separated "Flectonotus" (Anura: Hemiphractidae), as revealed by molecular, behavioral, and morphological data. Phyllomedusa: Journal of Herpetology, 10(1), 15-29. https://doi.org/10.11606/issn.2316-9079.v10i1p15-29