Lindquist, Erik, Michael Redmer, and Emily Brantner. 2012. “Annular Bone Growth in Phalanges of Five Neotropical Harlequin Frogs (Anura: Bufonidae: Atelopus)”. Phyllomedusa: Journal of Herpetology 11 (2): 117-24. https://doi.org/10.11606/issn.2316-9079.v11i2p117-124.