Barrio-Amorós, C.L., Santos, J.C. and Molina, C.R. (2010) “An addition to the diversity of dendrobatid frogs in Venezuela: description of three new collared frogs (Anura: Dendrobatidae: Mannophryne)”, Phyllomedusa: Journal of Herpetology, 9(1), pp. 03–35. doi:10.11606/issn.2316-9079.v9i1p03-35.