Barrio-Amorós, César Luis, et al. “An Addition to the Diversity of Dendrobatid Frogs in Venezuela: Description of Three New Collared Frogs (Anura: Dendrobatidae: Mannophryne)”. Phyllomedusa: Journal of Herpetology, vol. 9, no. 1, July 2010, pp. 03-35, https://doi.org/10.11606/issn.2316-9079.v9i1p03-35.